According to Principle II of the current International Code of Nomenclature for Algae, Fungi, and Plants (Madrid Code; hereinafter “the ICN”; Turland et al., 2025), the application of names of taxonomic groups is determined by means of nomenclatural types. In addition, Article 40.5 of the ICN (Turland et al., 2025) states that, “For the name of a new species or infraspecific taxon published on or after 1 January 1990 of which the type is a specimen or unpublished illustration, the single herbarium, collection, or institution in which the type is conserved must be specified.” Therefore, the type specimens of the species and the infraspecific taxon are important for taxonomic research, and it is necessary for researchers to examine the type specimen deposited in the specified herbarium, collection, or institution in order to apply the correct name for the taxon with a particular circumscription, position, and rank. Type specimens, however, can be transferred from a herbarium to another for a variety of reasons, resulting in difficulties for researchers examining the type specimens. Previously, Jang et al. (2020, 2022) provided information on more than 200 sheets of type specimens of vascular plants, most of which were donated from other herbaria to the herbarium of the National Institute of Biological Resources (KB).

Yong No Lee (1920–2008) played a key role in the history of Korean plant taxonomy and proposed 363 names of vascular plants as nomenclatural novelties, such as names of new taxa and new combinations through massive studies of the flora of the Korean Peninsula (Jang and Lee, 2022). He deposited most specimens, including types of names of new species and infraspecific taxa published by himself, in his personal but publicly accessible herbarium, the herbarium of the Korea Plant Research Institute (hereinafter “KPRI”). After he passed away on 22 June 2008, all specimens (approximately 12,000), including type specimens deposited at KPRI, were donated to KB by his family on 2 June 2009. In addition, nearly 60,000 slide films used for publishing names as well as his book, the New Flora of Korea (Lee, 2006), were transferred to KB at that time.

Nevertheless, information on Lee’s donated type specimens in KB remains not well documented, making it challenging to apply the names of species and infraspecific taxa published by Y. N. Lee. For example, Lee (2004a) indicated in the protologue of Veronica pusanensis Y. N. Lee that the type of this name was deposited into KPRI. After the donation of specimens to KB from KPRI, researchers did not find any specimens belonging to the original material of V. pusanensis, and Jang and Noh (2020) designated a neotype of the name. Upon our more carefully investigations, however, four specimens of the original material of the name were found in Lee’s donated collection in KB (see 40. V. pusanensis).

In order to ensure clarity and stability with regard to the taxonomic application of names of Korean species and infraspecific taxa, the type specimens and other elements of the original materials of the names published by Y. N. Lee were consulted by comparing all specimens in Lee’s donated collection in KB to information in the corresponding protologues. Information on the type specimens in Yong No Lee’s donated collection in KB is provided here. We and other members of KB are still searching for and examining type specimens and other elements of original materials of names; this information is not provided in this paper though it will be documented in the future.

The names of new species and infraspecific taxa validly published by Yong No Lee and the corresponding protologues were gathered. The nomenclatural statuses and the bibliographic references of the names were sourced from Jang and Lee (2022). In this study, names published only with indications in the corresponding protologues for which the types had been deposited in KPRI were investigated; these type specimens were donated to KB from KPRI after Lee’s death. By comparing all specimens in Yong No Lee’s donated collection in KB to information in the protologues, including descriptions and/or diagnoses, type citations, and illustrations, the type specimens and other original materials for the names were consulted and categorized in accordance with Article 9 of the ICN (Turland et al., 2025). Information regarding the type specimens is documented here, specifically the family, scientific name with author(s), bibliographic reference to valid publication, the Korean vernacular name, the type citation in the protologue (Protologue) for comparison with information on the type specimen, and details pertaining to the type specimen (Type). Current treatments of the name (i.e., whether the name was used as a correct one, as a homotypic synonym, or as a heterotypic synonym) by the National Institute of Biological Resources (hereinafter “NIBR,” 2019), Korea National Arboretum (hereinafter “KNA,” 2020), and according to Son et al. (2025) are also provided (Current). The names of species and infraspecific taxa were assigned to families according to NIBR (2019), and the families and the names of the species and infraspecific taxa were then arranged in alphabetical order.

All specimens, numbering approximately 12,000, in Yong No Lee’s donated collection in KB were examined and type specimens and other elements belonging to the original materials of the names published as names of new species and infraspecific taxa by Lee were consulted. In this study, only names validly published with citation for which types were deposited in KPRI were investigated. During the examination, we became aware that many specimens in the collection lack scientific names, Korean vernacular names, collection localities, collection dates, and/or the collector’s names. In addition, a number of consulted specimens appearing to be holotypes, syntypes, or even specimens clearly indicated as “type” on their labels contain different information from the type citations in the protologues. We also have come to presume strongly that in most cases Lee’s type citation in the protologue of a name was not based on a specimen itself but on information about when, where, or by whom plants of the corresponding taxon were discovered or their photographs in the protologue were taken. Therefore, it was challenging to find and categorize type specimens of names. For example, two specimens of Paspalum hallasanense Y. N. Lee (NIBRVP183360, 183361) were indicated as “Type” on their labels in Yong No Lee’s handwriting, and their collection locality and collection date match the type citation in the protologue of the name, but the name of the collector of the specimens differs from that of type citation in the protologue (see 35. P. hallasanense for details). It appears that they are syntypes of P. hallasanense and that the collector’s name was erroneously cited in the protologue. The specimens, however, are not syntypes and must be treated as uncited specimens of the original material of the name, as “Any designation of the type made by the original author(s), if definitely expressed in the protologue, is final” (Article 9.2 of the ICN; Turland et al., 2025) and because the current ICN allows corrections of errors made in the designation of a holotype, lectotype, neotype, or epitype, but not in the designation of a syntype (see Article 9.24 of the ICN; Turland et al., 2025) and the two specimens were not cited in the protologue of the name but were associated with the taxon by and available to the publishing author prior to publication of the protologue (see Article 9.4(e) and Article 9 Note 7 of The ICN; Turland et al., 2025).

As a result of evaluating and categorizing type specimens in Y. N. Lee’s donated collection in KB, we provide here information on 94 type specimens of 45 names validly published by Y. N. Lee. The type specimens consist of 11 holotypes (Fig. 1), 34 lectotypes (Figs. 2–4), and 49 isolectotypes. In addition, 45 names of 20 species, 11 varieties, and 14 forms were assignable to 27 genera of 23 families. The lectotypes of 34 names are designated here; if a holotype of a name was not found in Yong No Lee’s donated collection in KB, we assumed that the holotype was lost or destroyed and designated a lectotype of the name from the remaining elements of the original material.

1. Strobilanthes oligantha f. alba Y. N. Lee, Bull. Korea Pl. Res. 4: 15, 2004. 흰방울꽃.

Protologue: “Jeju-do, Aug. 12, 2004, Shin Yong Man & Moon Soon Hwa.”

Type: [KOREA.] Jeju-do: 12 Aug 2004, Soon Hwa Moon s.n. [lectotype, designated here: KB (NIBRVP193808!, Fig. 3A); isolectotype: KB (NIBRVP193810!)].

Note: Two specimens (NIBRVP193808, 193810) of Strobilanthes oligantha f. alba Y. N. Lee collected from Jeju-do on 12 August 2004 were consulted in Y. N. Lee’s donated collection in KB. The collector’s name of these specimens, however, differs from that of the type citation in the protologue; therefore, these two are not syntypes but are instead uncited specimens of the original material of S. oligantha f. alba. We did not find any other specimen of the original material for the name in Lee’s donated collection in KB and designate here NIBRVP193808 (Fig. 3A) as the lectotype; white corollas, the diagnostic morphological features of this taxon, are more clearly shown in this specimen.

Current: Heterotypic synonym of Strobilanthes oligantha Miq. (방울꽃, Son et al., 2025). Not treated by NIBR (2019) and KNA (2020).

2. Arisaema robustum f. variegatum Y. N. Lee, Bull. Korea Pl. Res. 5: 36, 2005. 무늬넓은천남성.

Protologue: “Mt. Mugap, Gyeonggi-do, Korea, May 5, 2004, Han Jin Su.”

Type: [KOREA.] Gyeonggi-do: Gwangju, Mt. Mugapsan, 5 May 2004, Yong No Lee s.n. [lectotype, designated here: KB (NIBRVP189674!, Fig. 3B); isolectotype: KB (NIBRVP189675!)].

Note: To KB from KPRI, three specimens (NIBRVP 189673–189675) of the genus Arisaema Mart. collected from Mt. Mugapsan on 5 May 2004 were donated. We carefully examined them and identified NIBRVP189674 and 189675 as A. robustum f. variegatum Y. N. Lee. However, the collector’s name for these two specimens does not match the protologue; thus, these specimens are uncited specimens of the original material of the name. We did not find any other elements of the original material in Lee’s donated collection in KB and designate here NIBRVP189674 (Fig. 3B) as the lectotype of A. robustum f. variegatum; a flower of another specimen, NIBRVP189675, has been damaged by insects.

Current: Heterotypic synonym of Arisaema amurense f. serratum (Nakai) Kitag. (천남성, Son et al., 2025). Not treated by NIBR (2019) and KNA (2020).

3. Asarum intermedium Y. N. Lee, Bull. Korea Pl. Res. 5: 5, 2005, non Grossh. 1930. 잡선운족도리풀.

Protologue: “Gochang, Jeollabuk-do, Korea, May 7, 2005, Lee Yong No & Kim You Sung.”

Type: [KOREA.] Jeollabuk-do: Gochang, 7 May 2005, Yong No Lee & You Sung Kim s.n. [holotype: KB (NIBRVP191440!, Fig. 1A)].

Current: Heterotypic synonym of Asarum maculatum Nakai (개족도리풀, Son et al., 2025). Not treated by NIBR (2019) and KNA (2020).

4. Asarum sieboldii var. cornutum Y. N. Lee, Bull. Korea Pl. Res. 1: 19, 2000. 뿔족도리풀.

Protologue: “Korea. Kangwon Province, Chindong Valley, May 22, 1998, Y. Lee.”

Type: [KOREA. Gangwon-do:] Jindong Valley, 22 May 1998, Yong No Lee s.n. [lectotype, designated here: KB (NIBRVP191454!, Fig. 2A); isolectotypes: KB (NIBRVP191450!–191453!)].

Note: Five syntypes (NIBRVP191450–191454) annotated as “Type: Asarum sieboldii Miquel var. cornutum Y. Lee” were donated to KB from KPRI. One specimen (NIBRVP191454, Fig. 2A) among them had an additional remark on its label “Type specimen,” and we designate here this specimen as the lectotype of A. sieboldii var. cornutum Y. N. Lee (see Recommendation 9A.3 of the ICN; Turland et al., 2025).

Current: Heterotypic synonym of Asarum sieboldii Miq. (족도 리풀, KNA, 2020; Son et al., 2025). Not treated by NIBR (2019).

5. Asarum sieboldii var. viridiluteolum Y. N. Lee, Bull. Korea Pl. Res. 5: 10, 2005. 황록족도리풀.

Protologue: “Mt. Baekdu, May 19, 2005, Lee Yong No & Kim You Sung.”

Type: [CHINA.] Mt. Baekdusan, 13–20 May 2005, Yong No Lee & You Sung Kim s.n. [lectotype, designated here: KB (NIBRVP190792!, Fig. 3C)].

Note: In Lee’s donated collection, we found three specimens (NIBRVP190790–92) with remarks on their labels as “Type: Asarum sieboldii Miquel var. viridiluteolum Y. Lee.” However, their collection dates differ from those in the type citation in the protologue; therefore, these are not syntypes but uncited specimens of the original material of A. sieboldii var. viridiluteolum Y. N. Lee, and the “Type” annotations on their labels are errors. One specimen (NIBRVP190792, Fig. 3C) was additionally annotated on its label as a “Type specimen,” and we designate here this specimen as the lectotype of the name (see Recommendation 9A.3 of the ICN; Turland et al., 2025). Because collection dates of the three specimens (NIBRVP190790–92) are not specific and as there is no evidence that they comprise a single gathering, we recognize that two specimens (NIBRVP190790 and 190791) are not duplicates of the lectotype (NIBRVP190792).

Current: Homotypic synonym of Asarum yeonbyeonense var. viridiluteolum (Y. N. Lee) M. Kim & S. So (녹연변족도 리풀, NIBR, 2019). Heterotypic synonym of A. yeonbyeonense M. Kim & S. So (연변족도리풀, KNA, 2020; Son et al., 2025).

6. Asarum sonunsanense Y. N. Lee, Bull. Korea Pl. Res. 5: 6, 2005. 선운족도리풀.

Protologue: “Gochang, Jeollabuk-do, Korea, May 7, 2005, Lee Yong No & Kim You Sung.”

Type: [KOREA.] Jeollabuk-do: Gochang, 7 May 2005, Yong No Lee & You Sung Kim s.n. [holotype: KB (NIBRVP208345!, Fig. 1B)].

Note: The red “TYPUS” annotation of the holotype (NIBRVP208345, Fig. 1B) of Asarum sonunsanense Y. N. Lee was attached by an anonymous member of KB in 2009 after the specimen had been donated to KB.

Current: Homotypic synonym of Asarum sieboldii var. sonunsanense (Y. N. Lee) M. Kim & S. So (KNA, 2020). Heterotypic synonym of A. maculatum Nakai (개족도리풀, Son et al., 2025). Not treated by NIBR (2019).

7. Asarum sonunsanense f. variegatum Y. N. Lee, Bull. Korea Pl. Res. 5: 7, 2005. 무늬선운족도리풀.

Protologue: “Gochang, Jeollabuk-do, Korea, May 7, 2005, Lee Yong No & Kim You Sung.”

Type: [KOREA. Jeollabuk-do:] Gochang, 7 May 2005, Yong No Lee & You Sung Kim s.n. [holotype: KB (NIBRVP191449!, Fig. 1C)].

Current: Heterotypic synonym of Asarum maculatum Nakai (개족도리풀, Son et al., 2025). Not treated by NIBR (2019) and KNA (2020).

8. Asarum sonunsanense f. variegatum Y. N. Lee, New Fl. Korea 2: 735, 2006, non Y. N. Lee 2005. 무늬황록선운족 도리풀.

Protologue: “Gochang, Jeollabuk-do, Korea, May 7, 2005, Lee Yong No & Kim You Sung.”

Type: [KOREA. Jeollabuk-do:] Gochang, 7 May 2005, Yong No Lee & You Sung Kim s.n. [lectotype, designated here: KB (NIBRVP191439!, Fig. 2B); isolectotype: KB (NIBRVP191438!)].

Note: Two syntypes (NIBRVP191438, 191439) of a later homonym Asarum sonunsanense f. variegatum Y. N. Lee were assessed in Lee’s donated collection in KB; they were indicated on their labels as “Type: Asarum sonunsanense Y. Lee var. viriluteolum (Y. Lee) Y. Lee for. variegatum Y. Lee.” On the label of NIBRVP191439 (Fig. 2B), the Korean words “무늬” and “황록색” were handwritten; these words indicate that a plant of the specimen has variegated (“무늬”) leaves and greenish yellow (“황록색”) calyces. This specimen (NIBRVP191439, Fig. 2B) is designated here as the lectotype of the name.

Current: Not treated by NIBR (2019), KNA (2020), and Son et al. (2025).

9. Cirsium setidens var. pinnatifolium Y. N. Lee, Bull. Korea Pl. Res. 4: 8, 2004. 깃잎고려엉겅퀴.

Protologue: “Mt. Hambaek, Aug. 31, 2003, Lee Yong No & Kim You Sung.”

Type: [KOREA. Gangwon-do:] Mt. Hambaeksan, 31 Aug 2003, You Sung Kim s.n. [lectotype, designated here: KB (NIBRVP190360!, Fig. 3D)].

Note: When validly publishing Cirsium setidens var. pinnatifolium Y. N. Lee, Lee and Kim (2004) distinguished the taxon from C. setidens (Dunn) Nakai var. setidens in having pinnate leaves. In Lee’s collection in KB, we found two specimens (NIBRVP190360, 190361) of C. setidens collected from Mt. Hambaeksan on 31 August 2003, and identified NIBRVP190360 (Fig. 3D) as C. setidens var. pinnatifolium and NIBRVP190361 as C. setidens var. setidens. However, the collector’s name for NIBRVP190360 (Fig. 3D) differs from that of the type citation in the protologue of C. setidens var. pinnatifolium; therefore, this specimen is not a holotype but is instead an uncited specimen of the original material of the name. We designate here NIBRVP190360 (Fig. 3D) as the lectotype of C. setidens var. pinnatifolium.

Current: Heterotypic synonym of Cirsium setidens (고려엉겅 퀴, KNA, 2020; Son et al., 2025). Not treated by NIBR (2019).

10. Impatiens kojeensis Y. N. Lee, Korean J. Pl. Taxon. 28: 28, 1998. 거제물봉선.

Protologue: “West south side of Isl. Kojedo, Kyongsangnamdo, Korea, Y. Lee, S. Cho and J. Kim s.n., Oct. 6, 1997.”

Type: [KOREA. Gyeongsangnam-do:] Isl. Geojedo, 6 Oct 1997, Yong No Lee s.n. [lectotype, designated here: KB (NIBRVP 186833!, Fig. 3E); isolectotypes: KB (NIBRVP186829!, 186830!, 186832!)].

Note: Four specimens (NIBRVP186829, 186830, 186832, 186833) identified as “거제물봉선” (Impatiens kojeensis Y. N. Lee) were donated to KB from KPRI. The collector’s name in this case, however, differs from that of the type citation in the protologue; thus, these specimens are uncited specimens of the original material of I. kojeensis. We did not find any other element of the original material for name in Lee’s donated collection in KB and designate here NIBRVP186833 (Fig. 3E) as the lectotype; this specimen is better preserved.

Current: Heterotypic synonym of Impatiens furcillata Hemsl. (처진물봉선, KNA, 2020; Son et al., 2025). Not treated by NIBR (2019).

11. Adenophora fusifolia Y. N. Lee, Bull. Korea Pl. Res. 4: 10, 2004. 백두모싯대.

Protologue: “Monte Baekdu, Aug. 1, 2004, Lee Yong No & Kim You Sung.”

Type: [CHINA.] Mt. Baekdusan, 1 Aug 2004, Yong No Lee s.n. [lectotype, designated here: KB (NIBRVP841592!, Fig. 3F); isolectotypes: KB (NIBRVP191475!, 191479!, 191482!–191484!)].

Note: Lee and Lee (2004a, 2004b) validly published Adenophora fusifolia Y. N. Lee and A. fusifolia f. albiflora Y. N. Lee based on materials collected by Yong No Lee and You Sung Kim from Mt. Baekdusan on 1 August 2004 and characterized plants having blue corollas as A. fusifolia f. fusifolia and those having white corollas as A. fusifolia f. albiflora. In Lee’s donated collection in KB, a total of ten specimens (NIBRVP191475–191479, 191482–191485, 841592) of A. fusifolia collected from Mt. Baekdusan on 1 August 2004 were consulted; all had their labels indicated as “중요 표본” (an important specimen). We carefully examined the color of the corollas and identified six (NIBRVP191475, 191479, 191482–191484, 841592) as A. fusifolia f. fusifolia and the others (NIBRVP191476–191478, 191485) as A. fusifolia f. albiflora (see 12. A. fusifolia f. albiflora).

Six specimens of A. fusifolia f. fusifolia (NIBRVP191475, 191479, 191482–191484, 841592) are not syntypes but are instead uncited specimens of the original material of the name, as the name of the collector of the specimens differs from that of the type citation in the protologue. Although one specimen among them, NIBRVP841592 (Fig. 3F), contained the remark “Type specimen” on its label and had the red “TYPUS” annotation, which was attached by an anonymous member of KB in 2009, it is not a type but an uncited specimen, and both the remark and the annotation are errors. We designate here the specimen NIBRVP841592 (Fig. 3F) as the lectotype of A. fusifolia.

Current: Correct name (백두모시대, KNA, 2020; Son et al., 2025). Not treated by NIBR (2019).

12. Adenophora fusifolia f. albiflora Y. N. Lee, Bull. Korea Pl. Res. 4: 13, 2004. 흰백두모싯대.

Protologue: “Mt. Baekdu, Aug. 1, 2004, Lee Yong No & Kim You Sung.”

Type: [CHINA.] Mt. Baekdusan, 1 Aug 2004, Yong No Lee s.n. [lectotype, designated here: KB (NIBRVP191478!, Fig. 3G); isolectotypes: KB (NIBRVP191476!, 191477!, 191485!)].

Note: As mentioned above (see 11. Adenophora fusifolia), we identified four specimens (NIBRVP191476–191478, 191485) as A. fusifolia f. albiflora. However, the name of the collector of the specimens differs from that on the type citation in the protologue; thus, these four specimens are uncited specimens composing the original material of A. fusifolia f. albiflora. We did not find any specimens that matched the type citation of the name in Lee’s donated collection; thus, we designate here NIBRVP191478 (Fig. 3G) as the lectotype. Although the lectotype was collected on 1 August 2004 and not on 3 August 2004 when a photograph of the taxon in the protologue was taken, it is very similar to the subject of the photograph in the morphology, such as the pattern of scratches on the leaves and the condition of four flowers: one flower is withered, another is closed, and the other two are fully opened (Fig. 3G).

Current: Heterotypic synonym of Adenophora fusifolia Y. N. Lee (백두모시대, Son et al., 2025). Not treated by NIBR (2019) and KNA (2020).

13. Dianthus chinensis f. albiflora Y. N. Lee, Bull. Korea Pl. Res. 4: 28, 2004. 흰난쟁이패랭이꽃.

Protologue: “Mt. Baekdu, Aug. 6, 2004, Lee Yong No & Kim You Sung.”

Type: [CHINA.] Mt. Baekdusan, Punggu, 6 Aug 2004, Yong No Lee & You Sung Kim s.n. [lectotype, designated here: KB (NIBRVP191489!, Fig. 2C); isolectotypes: KB (NIBRVP 191488!, 191490!)].

Note: Three syntypes (NIBRVP191488–191490) of Dianthus chinensis f. albiflora Y. N. Lee were donated to KB from KPRI; they were identified as “흰난쟁이패랭이꽃” (D. chinensis f. albiflora) on their labels. We designate here NIBRVP191489 (Fig. 2C) as the lectotype; white petals, the diagnostic morphological features of the taxon, are more clearly visible in this specimen.

Current: Heterotypic synonym of Dianthus chinensis L. (패랭이꽃, Son et al., 2025). Not treated by NIBR (2019) and KNA (2020).

14. Euonymus sieboldianus f. albicarpus Y. N. Lee, Bull. Korea Pl. Res. 5: 12, 2005. 흰참빗살나무.

Protologue: “Cheonan, Chungcheongnam-do, Korea, Nov. 9, 1998, Lee Yong No.”

Type: [KOREA. Chungcheongnam-do:] Cheonan, 9 Nov 1998, Yong No Lee s.n. [lectotype, designated here: KB (NIBRVP193078!, Fig. 2D); isolectotypes: KB (NIBRVP193075!–193077!)].

Note: We identified four specimens (NIBRVP193075–193078) in the donated collection in KB as syntypes of Euonymus sieboldianus f. albicarpus Y. N. Lee. Because white capsules are better observed in NIBRVP193078 (Fig. 2D), we designate here this specimen as the lectotype.

Current: Heterotypic synonym of Euonymus hamiltonianus Wall. (참빗살나무, Son et al., 2025). Not treated by NIBR (2019) and KNA (2020).

15. Orostachys chongsunensis Y. N. Lee, Bull. Korea Pl. Res. 1: 36, 2000. 정선바위솔.

Protologue: “Korea. Kangwon Province, Chongsun, Oct. 15, 1999, Y. Lee.”

Type: [KOREA. Gangwon-do:] Cheongseon, 15 Oct 1999, Yong No Lee 2 [holotype: KB (NIBRVP191442!, Fig. 1D)].

Current: Correct name (NIBR, 2019; KNA, 2020; Son et al., 2025).

16. Orostachys latielliptica Y. N. Lee, Bull. Korea Pl. Res. 1: 35, 2000. 포천바위솔.

Protologue: “Korea. Kyunggi Province, Pochon, Oct. 14, 1999, Y. Lee.”

Type: [KOREA. Gyeonggi-do:] Pocheon, 14 Oct 1999, Yong No Lee 3 [holotype: KB (NIBRVP191445!, Fig. 1E)].

Current: Correct name (NIBR, 2019; KNA, 2020; Son et al., 2025).

17. Orostachys ramosa Y. N. Lee, Bull. Korea Pl. Res. 1: 35, 2000. 가지바위솔.

Protologue: “Korea. Chunnam Province, Bosung, Nov. 23, 1999, Y. Lee.”

Type: [KOREA. Jeollanam-do:] Boseong, 23 Nov 1999, Yong No Lee s.n. [lectotype, designated here: KB (NIBRVP183933!, Fig. 2E)]; isolectotypes: KB (NIBRVP183930!, 183935!–183937!, 191444!)].

Note: Six syntypes (NIBRVP183930, 183933, 183935–183937, 191444) of Orostachys ramosa Y. N. Lee were assessed in Lee’s donated collection in KB. We designate here NIBRVP183933 (Fig. 2E) as the lectotype of the name; the diagnostic characteristics of this taxon, many-branched stems and obovate petals, are more clearly shown in this specimen.

Current: Correct name (NIBR, 2019; KNA, 2020; Son et al., 2025).

18. Swertia wilfordii f. albiflora Y. N. Lee, Bull. Korea Pl. Res. 5: 20, 2005. 흰큰잎쓴풀.

Protologue: “Mt. Jeombong, Gangwon-do, Korea, Sept. 14, 2002, Lee Yong No & Kim You Sung.”

Type: [KOREA.] Gangwon-do: Mt. Jeombongsan, 14–15 Sep 2002, Yong No Lee & You Sung Kim s.n. [lectotype, designated here: KB (NIBRVP185649!, Fig. 3H)].

Note: In Lee’s collection in KB, we found eight specimens (NIBRVP185641, 185642, 185644–185649) of Swertia wilfordii f. albiflora Y. N. Lee which were collected from Mt. Jeombongsan by Yong No Lee and You Sung Kim. However, the collection date for these specimens does not match the protologue; therefore, these specimens are uncited specimens of the original material of the name. We found no other elements of the original material in Lee’s donated collection in KB and designate here NIBRVP185649 (Fig. 3H) as the lectotype; white corollas are more visible in this specimen. We recognize that the other seven specimens (NIBRVP185641, 185642, 185644–185648) are not duplicates of the lectotype (NIBRVP185649), as the corresponding collection dates were not specified and because there is no evidence that they comprise a single gathering.

Current: Heterotypic synonym of Swertia wilfordii (A. Kern.) Kom. (큰잎쓴풀, Son et al., 2025). Not treated by NIBR (2019) and KNA (2020).

19. Iris neoensata Y. N. Lee, Bull. Korea Pl. Res. 5: 30, 2005. 신꽃창포.

Protologue: “Yongin, Gyeonggi-do, Korea, June 11, 2005, Lee Yong No.”

Type: [KOREA. Gyeonggi-do:] Yongin, 11 Jun 2005, Yong No Lee & You Sung Kim s.n. [lectotype, designated here: KB (NIBRVP189842!, Fig. 3I); isolectotype: KB (NIBRVP189844!)].

Note: When validly publishing Iris neoensata Y. N. Lee, Lee (2005) distinguished this taxon from I. ensata Thunb. in having narrow and long leaves and bracts. In Lee’s donated collection in KB, we found two specimens (NIBRVP189842, 189844) of I. neoensata collected from Yongin on 11 June 2005, and one of them, NIBRVP189842 (Fig. 3I), was identified on their label as “가는꽃창포” (narrow I. ensata). In addition, they appear to be the very plants photographed in the protologue of I. neoensata. However, the collector’s name for these two specimens differs from that of the type citation in the protologue; therefore, these two are uncited specimens composing the original material of I. neoensata. No other specimens matching the type citation in the protologue were found in the Lee’s donated collection in KB, and NIBRVP189842 (Fig. 3I) is designated here as the lectotype of the name; a flower of the other specimen (NIBRVP189844) is withered.

Current: Correct name (KNA, 2020; Son et al., 2025). Not treated by NIBR (2019).

20. Iris sanguinea var. coronalis Y. N. Lee, Bull. Korea Pl. Res. 5: 32, 2005. 한국붓꽃.

Protologue: “Mt. Odae, Gangwon-do, Korea, June 14, 2005, Lee Yong No & Kim You Sung.”

Type: [KOREA. Gangwon-do:] Mt. Odaesan, Botanical Garden of Korea Native Plant, 14 Jun 2005, Yong No Lee & You Sung Kim s.n. [holotype: KB (NIBRVP189843!, Fig. 1F)].

Current: Heterotypic synonym of Iris sanguinea Donn ex Hornem. (붓꽃, KNA, 2020; Son et al., 2025). Not treated by NIBR (2019).

21. Elsholtzia hallasanensis Y. N. Lee, Alp. Fl. Korea 328, 2000. 한라꽃향유.

Protologue: “Korea. Cheju Province, Cheju Isl. Mt. Halla, Nov. 4, 1998, Y. Lee & K. Lee.”

Type: [KOREA.] Jeju-do: 2–3 Nov 1998, Yong No Lee s.n. [lectotype, designated here: KB (NIBRVP194739!, Fig. 3J)].

Note: Lee (2000) indicated that the type of Elsholtzia hallasanensis Y. N. Lee was collected on 4 Nov 1998 from Mt. Hallasan in Jeju-do (Cheju Isl.). None of the specimens of the genus Elsholtzia Willd. collected on that date were assessed in Lee’s donated collection in KB, but there are four specimens (NIBRVP194735, 194736, 194739, 194742) of the genus collected from Jeju-do on 2–3 Nov 1998. We carefully examined these specimens and identified two (NIBRVP 194735, 194739) with purple corollas as E. hallasanensis f. hallasanensis and the others (NIBRVP194736, 194742) with white corollas as E. hallasanensis f. albiflora Y. N. Lee (see 22. E. hallasanensis f. albiflora). The former two (NIBRVP 194735, 194739) are uncited specimens of the original material of E. hallasanensis f. hallasanensis. The specimen NIBRVP194739 (Fig. 3J) is better preserved and is designated here as the lectotype of E. hallasanensis. Because the collection dates of two specimens (NIBRVP194735, 194739) of E. hallasanensis f. hallasanensis were not specified, we did not treat the two specimens as duplicates.

Current: Homotypic synonym of Elsholtzia splendens var. hallasanensis (Y. N. Lee) M. Kim (KNA, 2020; Son et al., 2025). Not treated by NIBR (2019).

22. Elsholtzia hallasanensis f. albiflora Y. N. Lee, Bull. Korea Pl. Res. 5: 53, 2005. 흰한라꽃향유.

Protologue: “Jeju-do, Korea, Nov. 2, 1999, Lee Yong No.”

Type: [KOREA.] Jeju-do: 2–3 Nov 1998, Yong No Lee s.n. [lectotype, designated here: KB (NIBRVP194736!, Fig. 3K)].

Note: As mentioned above (see 21. Elsholtzia hallasanensis), we identified two specimens (NIBRVP191736, 194742) having white corollas as E. hallasanensis f. albiflora. The collection date for these specimens differs from that of the type citation in the protologue; therefore, they are uncited specimens of the original material of E. hallasanensis f. albiflora. We designate here NIBRVP194736 (Fig. 3K) as the lectotype of the name; a plant on another specimen (NIBRVP194742) is in poor condition with few leaves and flowers. The specimen NIBRVP19742 is not an isolectotype, as the collection date of this specimen and the lectotype are not specific and because there is no evidence that they comprise a single gathering.

Current: Heterotypic synonym of Elsholtzia splendens var. hallasanensis (한라꽃향유, KNA, 2020; Son et al., 2025). Not treated by NIBR (2019).

23. Elsholtzia splendens f. roseola Y. N. Lee, Bull. Korea Pl. Res. 1: 50, 2000. 분홍꽃향유.

Protologue: “Korea. Kangwon Province, Mt. Chiak, Oct 15, 1999, Y. Lee.”

Type: [KOREA. Gangwon-do:] Mt. Chiaksan, 5 Oct 1999, Yong No Lee s.n. [lectotype, designated here: KB (NIBRVP183801, Fig. 3L)].

Note: Two specimens (NIBRVP183800, 183801) of Elsholtzia splendens Nakai ex F. Maek. collected from Mt. Chiaksan by Yong No Lee were assessed in Lee’s donated collection in KB. We identified NIBRVP183800 with reddish corollas as E. splendens f. splendens and NIBRVP183801 (Fig. 3L) with pinkish corollas as E. splendens f. roseola Y. N. Lee. The collection date of the latter specimen differs from that of the type citation in the protologue of E. splendens f. roseola; therefore, NIBRVP183801 (Fig. 3L) is an uncited specimen of the original material and is designated here as the lectotype of E. splendens f. roseola.

Current: Heterotypic synonym of Elsholtzia splendens Nakai ex F. Maek. (꽃향유, KNA, 2020; Son et al., 2025). Not treated by NIBR (2019).

24. Lilium amabile var. flavum Y. N. Lee, Fl. Korea, ed. 5, 1169, 2002. 노랑털중나리.

Protologue: “Korea. Kangwon Province, Mt. Taebaek, Jun. 9, 2002, Y. Kim.”

Type: [KOREA. Gangwon-do:] Mt. Daedeoksan, 19 Jun 2002, without collector’s name [lectotype, designated here: KB (NIBRVP191500!, Fig. 3M)].

Note: When Lilium amabile var. flavum Y. N. Lee was validly published, Lee (2002a) indicated that its type was collected from Mt. Taebaeksan on 9 June 2002. In the protologue, however, the distribution of the taxon is stated in Korean as “대덕산” (Mt. Daedeoksan) and not Mt. Taebaeksan. In addition, the photograph for the taxon in the protologue was taken from Mt. Daedeoksan and not Mt. Taebaeksan. After the name was validly published, Lee (2002b) published the same name again with a Latin description (see Jang and Lee, 2022) and stated that the type of L. amabile var. flavum was collected from Mt. Daedeoksan and not Mt. Taebaeksan; therefore, it appears that the type regarding the name was collected not from Mt. Taebaeksan but from Mt. Daedeoksan.

In Lee’s donated collection in KB, one specimen (NIBRVP191500, Fig. 3M) of Lilium amabile var. flavum was consulted. This specimen was collected from Mt. Daedeoksan but the collection date and collector’s name do not match the protologue for the name; therefore, this specimen is not a holotype but an uncited specimen of the original material of the name. We searched in vain for any other specimens of the original materials in Lee’s collection in KB and designate here NIBRVP191500 (Fig. 3M) as the lectotype of L. amabile var. flavum.

Current: Correct name (NIBR, 2019). Heterotypic synonym of Lilium amabile Palib. (털중나리, KNA, 2020; Son et al., 2025).

25. Lythrum salicaria f. albiflora Y. N. Lee, Bull. Korea Pl. Res. 5: 18, 2005. 흰털부처꽃.

Protologue: “Yongin, Gyeonggi-do, Korea, July 9, 2004, Lee Yong No & Kim You Sung.”

Type: [KOREA. Gyeonggi-do:] Yongin, 9 Jul 2004, without collector’s name [lectotype, designated here: KB (NIBRVP193705!, Fig. 3N); isolectotype: KB (NIBRVP193706!)].

Note: Two specimens (NIBRVP193705, 193706) of Lythrum salicaria f. albiflora Y. N. Lee, the collection locality and date of which match the protologue, were donated to KB from KPRI. However, the collector’s name was not written on the corresponding labels; therefore, these specimens are uncited specimens of the original material of L. salicaria f. albiflora. We failed to find any other specimen of the original material of the name in Lee’s donated collection in KB and designate here NIBRVP193705 (Fig. 3N) as the lectotype; this specimen is in better condition than the other.

Current: Heterotypic synonym of Lythrum salicaria L. (털 부처꽃, Son et al., 2025). Not treated by NIBR (2019) and KNA (2020).

26. Cypripedium agnicapitatum Y. N. Lee, Fl. Korea, ed. 5, 1170, 2002. 양머리복주머니란.

Protologue: “China. Mt. Jangbae, June 6, 2002, Y. Lee & Y. Kim.”

Type: [CHINA. Mt. Changbai,] Hwaryong, 6 Jun 2002, Yong No Lee & You Sung Kim s.n. [lectotype, designated here: KB (NIBRVP185175!, Fig. 2F); isolectotypes: KB (NIBRVP 185174!, 185176!, 185177!)].

Note: Four syntypes (NIBRVP185174–185177) of Cypripedium agnicapitatum Y. N. Lee were donated to KB from KPRI. Three (NIBRVP185174, 185176, 185177) were identified on their labels as “양머리복주머니란” (C. agnicapitatum), but the other (NIBRVP185175) was identified as “양두복주머니”; both “머리” and “두” have the same meaning: “a head.” We designate here the specimen NIBRVP185175 (Fig. 2F) as the lectotype of the name; the specimen is better preserved.

Current: Correct name (KNA, 2020; Son et al., 2025). Not treated by NIBR (2019).

27. Cypripedium guttatum f. punicum Y. N. Lee, Bull. Korea Pl. Res. 5: 63, 2005. 분홍털복주머니란.

Protologue: “Mt. Baekdu, June 26, 2001, Lee Yong No.”

Type: [CHINA.] Mt. Baekdusan, 28 Jun 2001, without collector’s name [lectotype, designated here: KB (NIBRVP208346!, Fig. 3O)].

Note: Only one specimen (NIBRVP208346, Fig. 3O) of Cypripedium guttatum f. punicum Y. N. Lee collected from Mt. Baekdusan in 2001 was donated to KB from KPRI. This specimen, however, is not a holotype but is instead an uncited specimen belonging to the original material, as both the collection date and collector’s name differ between the label of the specimen and the protologue of the name. We designate here this specimen (NIBRVP208346, Fig. 3O) as the lectotype of C. guttatum f. punicum.

Current: Heterotypic synonym of Cypripedium guttatum Sw. (털복주머니란, KNA, 2020; Son et al., 2025). Not treated by NIBR (2019).

28. Cypripedium morinanthum Y. N. Lee, Fl. Korea, ed. 5, 1170, 2002. 자주복주머니란.

Protologue: “China. Mt. Jangbae, June 1, 2002, Y. Lee & Y. Kim.”

Type: [CHINA. Mt. Changbai,] Samdo, 1 Jun 2002, Yong No Lee & You Sung Kim s.n. [lectotype, designated here: KB (NIBRVP185129!, Fig. 2G); isolectotype: KB (NIBRVP185122!)].

Note: When two syntypes (NIBRVP185122, 185129) of Cypripedium morinanthum Y. N. Lee were donated to KB from KPRI, they were wrapped together in a single newspaper on which their flower color, collection locality, and collection date were written in Chinese characters (Fig. 2G). In addition, they were identified as “자주복주머니란” (C. morinanthum) in Yong No Lee’s handwriting. We designate here NIBRVP185129 (Fig. 2G) as the lectotype; a flower of the specimen is better preserved.

Current: Heterotypic synonym of Cypripedium× ventricosum Sw. (얼치기복주머니란, KNA, 2020; Son et al., 2025). Not treated by NIBR (2019).

29. Cypripedium neoparviflorum Y. N. Lee, Fl. Korea, ed. 5, 1171, 2002. 미색복주머니란.

Protologue: “China. Samdo, Mt. Baetoo, June 9, 2002, Y. Lee & Y. Kim.”

Type: [CHINA. Mt. Baekdusan,] Samdo, 9 Jun 2002, Yong No Lee & You Sung Kim s.n. [lectotype, designated here: KB (NIBRVP185132!, Fig. 2H); isolectotypes: KB (NIBRVP185133!, 185134!)].

Note: Three syntypes (NIBRVP185132–185134) of Cypripedium neoparviflorum Y. N. Lee were identified on their label as “백색복주머니란” (for example, see Fig. 2H) and not “미색복주머니란;” the meaning of “백색” (white color) is similar to that of “미색” (rice color). Because NIBRVP185132 (Fig. 2H) is in better condition, we designate here this specimen as the lectotype.

Current: Heterotypic synonym of Cypripedium macranthos Sw. (복주머니란, KNA, 2020; Son et al., 2025). Not treated by NIBR (2019).

30. Cypripedium sinapoides Y. N. Lee, Bull. Korea Pl. Res. 2: 30, 2002. 겨자복주머니란.

Protologue: “China. Mt. Baektu (Mt. Jangbaek), June 2, 2002, Y. Kim & Y. Lee.”

Type: [CHINA.] near Sinhap, 8 Jun 2002, Yong No Lee & You Sung Kim s.n. [lectotype, designated here: KB (NIBRVP185114!, Fig. 3P)].

Note: Three specimens (NIBRVP185112–185114) identified as “겨자복주머니란” (Cypripedium sinapoides Y. N. Lee) in Yong No Lee’s handwriting were assessed in Lee’s donated collection in KB. Their collection dates, however, did not match the type citation in the protologue of C. sinapoides; NIBRVP185112 was collected on 5 June 2022, NIBRVP185113 on 1–10 June 2002, and NIBRVP185114 (Fig. 3P) on 8 June 2002. Therefore, they are uncited specimens of the original materials of the name. We designate here the specimen NIBRVP185114 (Fig. 3P) as the lectotype of C. sinapoides; its collection date is more specific and the color of the perianth is denoted on its label as “겨자색” (mustard color).

Current: Heterotypic synonym of Cypripedium× ventricosum Sw. (얼치기복주머니란, KNA, 2020; Son et al., 2025). Not treated by NIBR (2019).

31. Goodyera rosulacea Y. N. Lee, Bull. Korea Pl. Res. 4: 2, 2004. 로젯사철란.

Protologue: “Mt. Dobong, June 18, 2002, Lee Yong No & Kim You Sung.”

Type: [KOREA.] Mt. Dobongsan, 18 Jun 2003, without collector’s name [lectotype, designated here: KB (NIBRVP191496!, Fig. 3Q)].

Note: One specimen (NIBRVP191496, Fig. 3Q) collected from Mt. Dobongsan and identified as “로젯사철란” (Goodyera rosulacea Y. N. Lee) in Yong No Lee’s handwriting was donated to KB from KPRI. This specimen, however, is not a holotype but is instead an uncited specimen of the original material, as the collection date and the collector’s name differ from those of the type citation in the protologue. We designate here NIBRVP191496 (Fig. 3Q) as the lectotype of G. rosulacea.

Current: Correct name (NIBR, 2019; 한국사철란, KNA, 2020). Heterotypic synonym of Goodyera brachystegia Hand.-Mazz. (로젯사철란, Son et al., 2025).

32. Hylomecon hylomeconoides var. dissectifolia Y. N. Lee, Bull. Korea Pl. Res. 4: 27, 2004. 갈래피나물.

Protologue: “Mt. Baekun, Jeollabuk-do, April 10, 2004, Lee Yong No & Kim Sun Wan.”

Type: [KOREA. Jeollanam-do:] Gurye, Mt. Baekunsan, 10 Apr 2004, without collector’s name [holotype: KB (NIBRVP191446!, Fig. 1G)].

Note: Lee (2004b) stated that the type of Hylomecon hylomeconoides var. dissectifolia Y. N. Lee was collected “in Mt. Baekunsan, Jeollabuk-do,” but Jeollabuk-do province contains at least three mountains called Mt. Baekunsan. In the protologue of the name, Lee (2004b) also denoted the collection locality of the type in Korean as “기준 표본: 전북 구례 백운산” (type specimen: Jeollabuk-do, Gurye, Mt. Baekunsan). However, Gurye (Gurye-gun, an administrative district in Korea) belongs to Jellanam-do province and not to Jellabuk-do; therefore, it appears that the type specimen was collected from Mt. Baekunsan in Gurye, Jeollanam-do.

In Lee’s collection in KB, we consulted only one specimen (NIBRVP191446, Fig. 1G) of H. hylomeconoides var. dissectifolia; it was collected from Gurye, Mt. Baekunsan, on 10 April 2004. Although no collector’s name is written on its label, this specimen was indicated as “Type specimen” (Fig. 1G); therefore, it is the holotype of H. hylomeconoides var. dissectifolia. The type citation in the protologue including the collection locality and collector’s name is erroneous and is to be corrected (see Article 9.24 of the ICN; Turland et al., 2025).

Current: Heterotypic synonym of Coreanomecon hylomeconoides Nakai (매미꽃, Son et al., 2025). Not treated by NIBR (2019) and KNA (2020).

33. Miscanthus latissimus Y. N. Lee, Bull. Korea Pl. Res. 6: 13, 2006. 장수억새.

Protologue: “Mt. Sobaek, Korea, Sept. 12, 2006, Lee Yong No & Kim You Sung.”

Type: [KOREA.] Mt. Sobaeksan, 12 Sep 2006, without collector’s name [holotype: KB (NIBRVP183383!, Fig. 1H)].

Note: Ibaragi et al. (2017) designated the specimen, K. I. Heo et al. 20110916028 (TKPM BSP-079819) as the neotype of Miscanthus latissimus Y. N. Lee, because at that time the depository of its holotype was quite unknown and no other specimen was known at all; when a request to KB to examine the type specimens of M. latissimus was made by Ibaragi et al. (2017), a member of KB did not assess any type specimens nor other elements of the original material. As a result of our careful investigation, only one specimen of M. latissimus (NIBRVP183383, Fig. 1H) denoted as “장수억새” (Miscanthus latissimus) and “기준표본” (a type specimen) on its label was assessed in Lee’s donated collection. (The red “TYPUS” annotation in this specimen, which is not original, was attached by an unknown member of KB in 2009.) Although no collector’s name is written on its label, the specimen (NIBRVP183383, Fig. 1H) is the holotype of M. latissimus. Corrected information on the holotype is provided here (see Article 9.24 of the ICN; Turland et al., 2025).

Current: Correct name (Son et al., 2025). Not treated by NIBR (2019) and KNA (2020).

34. Miscanthus sinensis var. viridis Y. N. Lee, Bull. Korea Pl. Res. 6: 16, 2006. 연두억새.

Protologue: “Pocheon, Gyeonggi-do, Korea, Sept. 22, 2006, Lee Yong No & Kim You Sung.”

Type: [KOREA. Gyeonggi-do: Pocheon,] Pyeonggang Botanical Garden, 22 Sep 2006, without collector’s name [holotype: KB (NIBRVP183382!, Fig. 1I)].

Note: Ibaragi et al. (2017) noted that no type specimens or any other elements of the original material for the name Miscanthus sinensis var. viridis Y. N. Lee had been found. During our examination of Lee’s donated collection in KB, one specimen (NIBRVP183382, Fig. 1I) of M. sinensis var. viridis collected on 22 Sep 2006 was consulted. The specimen was collected from the Pyeonggang Botanical Garden and its collector’s name was not written on its label. Nevertheless, the label of this specimen was denoted as “연두억새” (M. sinensis var. viridis) and “기준표본” (a type specimen), and the Pyeonggang Botanical Garden is located in Pocheon; therefore, this specimen (NIBRVP183382, Fig. 1I) is the holotype of M. sinensis var. viridis. We provide here corrected information on the holotype (see Article 9.24 of the ICN; Turland et al., 2025).

Current: Heterotypic synonym of Miscanthus sinensis Andersson (참억새, KNA, 2020; Son et al., 2025). Not treated by NIBR (2019).

35. Paspalum hallasanense Y. N. Lee, Bull. Korea Pl. Res. 7: 10, 2007. 한라참새피.

Protologue: “Jeju-do, Korea, Sep. 29, 2007, Lee Yong No.”

Type: [KOREA.] Jeju-do: 29 Sep 2007, without collector’s name [lectotype, designated here: KB (NIBRVP183360!, Fig. 3R); isolectotype: KB (NIBRVP183361!)].

Note: In Lee’s collection in KB, two specimens (NIBRVP183360, 183361) identified as “Paspalum hallasanense Y. Lee” in Yong No Lee’s handwriting were assessed; their collection locality and date match the type citation in the protologue of P. hallasanense Y. N. Lee. Although these specimens were annotated as “Type” in Yong No Lee’s handwriting, the collector’s name differs from that of the type citation in the protologue; therefore, they are not syntypes but are instead uncited specimens belonging to the original material of the name. Because a few spikelets remain on the NIBRVP183361, we designate here NIBRVP183360 (Fig. 3R) as the lectotype.

Current: Not treated by NIBR (2019), KNA (2020), and Son et al. (2025).

36. Hepatica asiatica var. yuseongii Y. N. Lee, Bull. Korea Pl. Res. 7: 8, 2007. 유성노루귀.

Protologue: “Pungdo, Gyeonggi-do, Korea, May 10, 2007, Kim You Sung.”

Type: [KOREA. Gyeonggi-do:] Isl. Pungdo, 9 Mar 2007, without collector’s name [lectotype, designated here: KB (only a material in the right pocket on NIBRVP183378!, Fig. 4A)].

Note: In the protologue of Hepatica asiatica var. yuseongii Y. N. Lee, Lee (2007) distinguished this taxon from H. asiatica Nakai var. asiatica in having linear perianths (petaloid sepals) and acuminate lobed leaves, indicating that its type was collected on 10 May 2007 from Isl. Pungdo. In the protologue, Lee (2007) also provided four photographs of the taxon, all of which were taken on 10 May 2007; two flowers of H. asiatica var. yuseongii are shown in the third photograph (Fig. 4B), and flowers of H. asiatica var. yuseongii and H. asiatica var. asiatica were taken for comparison and are shown in the fourth (Fig. 4C).

One specimen (NIBRVP183378, Fig. 4A) with its label denoted as “노루귀 ( 가는 잎)” [H. asiatica (narrow leaves)] and “중요기준표본” (an important type specimen) was assessed in Lee’s donated collection in KB; this specimen was collected on 9 March 2007 and not on 10 May 2007. On this specimen, two pockets containing flowers are mounted (Fig. 4A); these pockets were also denoted as “중요기준표본” (an important type specimen). The right pocket of the specimen contains a flower of H. asiatica var. yuseongii (Fig. 4A); this flower appears to be the one on the left in the third photograph in the protologue (Fig. 4B). The left pocket of the specimen contains two flowers; one is of H. asiatica var. asiatica and the other is of H. asiatica var. yuseongii (Fig. 4A), and it appears that they are the subjects of the fourth photograph in the protologue (Fig. 4C). From Lee’s donated slide films, we assessed the original films of the third (Fig. 4D) and the fourth photograph (Fig. 4E) used for the protologue of H. asiatica var. yuseongii. These two films were taken on 10 March 2007 and not on 10 May 2007, and the date of photoshoots does not match the protologue; therefore, it appears that the date on which photographs were taken was erroneously recorded in the protologue of H. asiatica var. yuseongii, and it is questionable as to whether the date on which the type specimen of the name was collected is 10 May 2007.

Because no other elements of the original material of H. asiatica var. yuseongii were found in Lee’s donated collection in KB and considering that the specimen NIBRVP183378 is annotated as “중요기준표본” (an important type specimen; Fig. 4A), we recognized that the specimen NIBRVP183378 (Fig. 4A) is the holotype of H. asiatica var. yuseongii and that the collection date and the collector’s name of the type citation in the protologue are erroneous (see Article 9.24 of the ICN; Turland et al., 2025). However, the materials on the holotype are mixed with a flower of H. asiatica var. asiatica in the left pocket (Fig. 4A). Thus, according to Article 9.11(c) of the ICN (Turland et al., 2025), we designate here the material in the right pocket on the specimen NIBRVP183378 (Fig. 4A) as the lectotype of H. asiatica var. yuseongii.

Current: Homotypic synonym of Hepatica asiatica f. yuseongii (Y. N. Lee) M. Kim (KNA, 2020; Son et al., 2025). Not treated by NIBR (2019).

37. Ranunculus franchetii var. hallasanensis Y. N. Lee, Bull. Korea Pl. Res. 7: 2, 2007. 한라왜미나리아재비.

Protologue: “Isl. Jeju, Korea, Mar. 27, 2007, Lee Yong No & Kim You Sung.”

Type: [KOREA.] Jeju-do: 24 Mar 2007, without collector’s name [holotype: KB (NIBRVP183379!, Fig. 1J)].

Note: In Lee’s donated collection in KB, only one specimen (NIBRVP183379, Fig. 1J) of Ranunculus franchetii var. hallasanensis Y. N. Lee was consulted. Although the collection date differs from that of type citation in the protologue, this specimen was denoted as “한라왜미나리아재비” (R. franchetii var. hallasanensis) and “기준표본” (type specimen) on its label; therefore, NIBRVP183379 (Fig. 1J) is the holotype of the name. The type citation in the protologue is an error and corrected information on the holotype is provided here (see Article 9.24 of the ICN; Turland et al., 2025).

Current: Heterotypic synonym of Ranunculus franchetii H. Boissieu (왜미나리아재비, KNA, 2020; Son et al., 2025). Not treated by NIBR (2019).

38. Duchesnea indica var. albicarpa Y. N. Lee, Bull. Korea Pl. Res. 4: 21, 2004. 흰뱀딸기.

Protologue: “in east part of Jeju-do, June 2, 2004, Lee Yong No & Shin Yong Man.”

Type: [KOREA.] Jeju-do: 2 Jun 2004, Yong No Lee s.n. [lectotype, designated here: KB (NIBRVP191073!, Fig. 3S); isolectotypes: KB (NIBRVP191071!, 191072!, 191074!)].

Note: Four specimens (NIBRVP191071–191074) identified as “흰뱀딸기” (Duchesnea indica var. albicarpa Y. N. Lee) on their labels were donated to KB from KPRI. These specimens are uncited specimens of the original material of D. indica var. albicarpa because the collector’s name differs from that of the type citation in the protologue. We failed to find any other specimen of the name in Lee’s donated collection in KB and designate here the specimen NIBRVP191073 (Fig. 3S) as the lectotype; the diagnostic features of the taxon, white receptacles and seeds, are more clearly shown in this specimen.

Current: Heterotypic synonym of Duchesnea indica (Andrews) Teschem. (뱀딸기, Son et al., 2025). Not treated by NIBR (2019) and KNA (2020).

39. Mazus pumilus f. albiflorus Y. N. Lee, Bull. Korea Pl. Res. 5: 21, 2005. 흰주름잎.

Protologue: “Busan, Korea, July 1, 2004, Lee Yong No & Kim You Sung.”

Type: [KOREA.] Busan: 1 Jul 2004, Yong No Lee & You Sung Kim s.n. [holotype: KB (NIBRVP191455!, Fig. 1K)].

Current: Heterotypic synonym of Mazus pumilus (Burm.f.) Steenis (주름잎, Son et al., 2025). Not treated by NIBR (2019) and KNA (2020).

40. Veronica pusanensis Y. N. Lee, Bull. Korea Pl. Res. 4: 16, 2004. 부산꼬리풀.

Protologue: “Wol Jeon, Busan, Oct. 2, 2004, Lee Yong No, Jeong Hyeon Do, Lee Bong Sik & Lee Mong Hwan.”

Type: [KOREA.] Busan: 2 Oct 2004, Yong No Lee & You Sung Kim s.n. [lectotype, designated here: KB (NIBRVP191503!, Fig. 3T); isolectotypes: KB (NIBRV P191502!, 191504!, 191505!)].

Note: Jang and Noh (2020) recognized that the holotype of Veronica pusanensis Y. N. Lee is missing and other elements of the original material of the name no longer exist, and they designated the specimen H. D. Jang 505 (KB, NIBRVP752966!) as the neotype of V. pusanensis. We, however, consulted four uncited specimens of the original material (NIBRVP191502–191505) regarding the name in Lee’s donated collection in KB. They contain different information about the collector’s names compared to the type citation in the protologue, but they were indicated as “중요표 본” (an important specimen) and identified as “Veronica pusanensis Y. Lee” in Yong No Lee’s handwriting. The specimen NIBRVP191503 (Fig. 3T) was the best preserved and is designated here as the lectotype, superseding the choice of the neotype by Jang and Noh (2020) in accordance with Article 9.19(a) of the ICN (Turland et al., 2025).

Current: Homotypic synonym of Pseudolysimachion pusanensis (Y. N. Lee) Y. N. Lee (NIBR, 2019; KNA, 2020; Son et al., 2025).

41. Scopolia kwangdokensis Y. N. Lee, Bull. Korea Pl. Res. 5: 22, 2005. 광덕미치광이풀.

Protologue: “Mt. Gwangdeok, Gangwon-do, Korea, April 26, 2005, Lee Yong No & Kim You Sung.”

Type: [KOREA. Gangwon-do:] Mt. Gwangdeoksan, 26 Apr 2005, Yong No Lee and You Sung Kim s.n. [lectotype, designated here: KB (NIBRVP187134!, Fig. 2I); isolectotypes: KB (NIBRVP 187135!, 187136!)].

Note: Three syntypes (NIBRVP187134–36) of Scopolia kwangdokensis Y. N. Lee were donated to KB from KPRI. One specimen, NIBRVP187134 (Fig. 2I), is designated here as the lectotype; brownish yellow corollas are more clearly shown in this specimen.

Current: Heterotypic synonym of Scopolia parviflora (Dunn) Nakai (미치광이풀, KNA, 2020; Son et al., 2025). Not treated by NIBR (2019).

42. Vitex rotundifolia f. albiflora Y. N. Lee, Bull. Korea Pl. Res. 5: 26, 2005, non S. S. Ying 1987. 흰순비기나무.

Protologue: “Daebudo, Gyeonggi-do, Korea, Aug. 22, 2005, Kim You Sung.”

Type: [KOREA.] Gyeonggi-do: Isl. Daebudo, 22 Aug 2005, You Sung Kim s.n. [lectotype, designated here: KB (NIBRVP 192579!, Fig. 2J); isolectotypes: KB (NIBRVP188485!, 188486!)].

Note: Three syntypes (NIBRVP188485, 188486, 192579) of a later homonym Vitex rotundifolia f. albiflora Y. N. Lee were assessed in Lee’s donated collection in KB. We designate here NIBRVP192579 (Fig. 2J), which obviously has white corollas, as the lectotype of the name.

Current: Heterotypic synonym of Vitex rotundifolia L. f. (순비기나무, Son et al., 2025). Not treated by NIBR (2019) and KNA (2020).

43. Viola albida var. rugata Y. N. Lee, Bull. Korea Pl. Res. 5: 16, 2005. 오골제비꽃.

Protologue: “Pocheon, Gyeonggi-do, Korea, May 6, 2005, Lee Yong No & Kim You Sung.”

Type: [KOREA. Gyeonggi-do:] Pocheon, Gaksibong, 5 May 2005, You Sung Kim s.n. [lectotype, designated here: KB (NIBRVP191493!, Fig. 3U); isolectotype: KB (NIBRVP191494!)].

Note: We identified two specimens (NIBRVP191493, 191494) collected in Pocheon by You Sung Kim as Viola albida var. rugata Y. N. Lee; they are indicated as “중요표본” (important specimen). The collection date of these specimens differs from that of the type citation in the protologue; therefore, these specimens are not the syntype but are instead uncited specimens belonging to the original material of V. albida var. rugata. We designate here NIBRVP191493 (Fig. 3U) as the lectotype of the name; the diagnostic features of the taxon, wrinkled flower petals and long triangular leaves, are more evidently shown in this specimen.

Current: Heterotypic synonym of Viola albida Palib. (태 백제비꽃, KNA, 2020; Son et al., 2025). Not treated by NIBR (2019).

44. Viola chejuensis Y. N. Lee & Y. C. Oh, Bull. Korea Pl. Res. 5: 13, 2005. 제주제비꽃

Protologue: “Sangumbul, Jeju-do, Korea, April 22, 2005, Y. Lee & Y. Oh.”

Type: [KOREA.] Jeju-do: 21–23 Apr 2005, Yong No Lee & You Sung Kim s.n. [lectotype, designated here: KB (NIBRVP189834!; Fig. 3V)].

Note: We consulted and identified only one specimen (NIBRVP189834; Fig. 3V) as Viola chejuensis Y. N. Lee & Y. C. Oh in Lee’s donated collection in KB. The collection date and the collector’s name of the specimens differ from those of the type citation in the protologue; therefore, this specimen is not a holotype but an uncited specimen of the original material concerning the name. We designate here NIBRVP189834 (Fig. 3V) as the lectotype.

Current: Transferred to the hybrid category, Viola × chejuensis (KNA, 2020; Son et al., 2025). Not treated by NIBR (2019).

45. Viola palatina Y. N. Lee, Bull. Korea Pl. Res. 5: 15, 2005. 창덕제비꽃.

Protologue: “Changdeok Palace, Seoul, Korea, April 5, 2005, Lee Yong No.”

Type: [KOREA. Seoul:] Changdeok Palace, 5 Apr 2005, Lee Yong No s.n. [lectotype, designated here: KB (NIBRVP189848!, Fig. 2K); isolectotype: KB (NIBRVP 189847!)].

Note: We consulted two syntypes (NIBRVP189847, 189848) of Viola palatina Y. N. Lee in Lee’s donated collection in KB, and designate here NIBRVP189848 (Fig. 2K) as the lectotype; this specimen was better preserved.

Current: Transferred to the hybrid category, Viola × palatina (KNA, 2020; Son et al., 2025). Not treated by NIBR (2019).